Although the Mamluks had taken over Cilicia, they were unable to hold it. Turkic tribes settled there, leading to the conquest of Cilicia led by Timur. As a result, 30,000 wealthy Armenians left Cilicia and settled in Cyprus, still ruled by the Lusignan dynasty until 1489. Many merchant families also fled westward and founded or joined with existing diaspora communities in France, Italy, the Netherlands, Poland, and Spain. Only the humbler Armenians remained in Cilicia. They nevertheless maintained their foothold in the region throughout Turkish rule.